Beatriz Arroyo

bea Beatriz Arroyo

Position: Científico Titular del CSIC
Address:  IREC .
Degree: Doctor.

I work at IREC since 2006. Before that, I spent four years in Oxford, UK, at the Edward Grey Institute of Field Ornithology (EGI, to carry out my PhD; four years in France at the Centre d´Etudes Biologiques de Chizé (CEBC-CNRS;), as a postdoctoral researcher, and six years in Scotland in the Center of Ecology and Hydrology, Banchory (CEH-Banchory;). My research has as the main objective to contribute to compatibilze human activities and wildlife conservation. Among others, my research interests include the following topics:

  • Ecology and conservation of harrier species (Montagus, pallid, hen and black harriers).
  • Agriculture and conservation of farmland birds.
  • Game management and environmental sustainability.
  • Effects of human disturbance on protected birds.
  • Management of ecological and conservation conflicts.

I am also increasingly interested by non-ecological (social and economic) aspects associated to nature conservation, and how these relate to transferability of research results to management decisions. Additionally, I am also interested in cost-efficiency and long-term sustainability of wildlife management and conservation.

Articles in SCI Journals||Other Articles

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Articles in SCI Journals

In press|2017|2016|2015|2014|2013|2012-1995

In press

  • Arroyo B., Mougeot F., Bretagnolle V. In press. Individual variation in behavioural responsiveness to humans leads to differences in breeding success and long-term population phenotypic changes. Ecology Letters. DOI: 10.1111/ele.12729
  • Garcia-Heras M-S, Mougeot, F., Arroyo, B., Avery, M., Simmons, R.E. In press. Is the Black Harrier Circus maurus a specialist predator? Assessing the diet of a threatened raptor species endemic to southern Africa. Ostrich. DOI: 10.2989/00306525.2016.1257515
  • Arroyo, A., Caro, J., Muñoz-Adalia, E.J., Díaz-Fernández, S., Delibes-Mateos, M., Díaz-Fernández, M., Viñuela, J. In press. Reconciling economic and ecological sustainability: Can non-intensive hunting of red-legged partridges be economically profitable? European Journal of Wildlife Research. DOI 10.1007/s10344-016-1073-2
  • Delibes-Mateos M, Caro J, Arroyo B. Accepted. Funciones y valoraciones de la caza y su gestión en España: estudios científicos sobre el colectivo cinegético. Arbor.

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  • Casas F, Arroyo B, Viñuela J, Guzmán JL, Mougeot F (2016) Are farm-reared red-legged partridge releases increasing hunting pressure on wild partridge breeding stocks? European Journal of Wildlife Research 62: 79-84
  • Díaz M, Moreno E, Amat JA, Arroyo B, Barba E, González-Solís J, Laiolo P, De Lope F, Merino S, Obeso JR & Velando A. (2016) Ardeola, a scientific journal of ornithology: cooperative survivorship within the red queen game. Ardeola 63: 3-14
  • Díaz-Ruiz, F., Caro, J., Delibes-Mateos, M., Arroyo, B. y Ferreras, P. (2016) Drivers of red fox (Vulpes vulpes) daily activity: prey availability, human disturbance or habitat structure? Journal of Zoology, 298: 128-138.
  • Estrada, A., Delgado, M-P., Arroyo, B., Traba, J. & Morales, M. (2016) Forecasting large-scale habitat suitability of European bustards under climate change: the role of environmental and geographic variables. PLOS One 11(3): e0149810
  • Garcia-Heras M.S., Arroyo B., Mougeot F., Amar A., Simmons R.E. (2016) Does timing of breeding matter less where the grass is greener? Seasonal declines in breeding performance differ between regions in an endangered endemic raptor. Nature Conservation 15: 23-45
  • Mills J.A., Teplitsky C., Arroyo B. Charmantier A., Becker P.H., Birkhead T.R., Bize P., Blumstein DT, Bonenfant C., Boutin S, Bushuev A., Cam E., Cockburn A., Côté SD, Coulson JC, Daunt F, Dingemanse NJ, Doligez B., Drummond H., Espie RHM, Festa-Bianchet M., Frentiu F., Fitzpatrick JW, Furness RW, Garant D., Gauthier G., Grant PR, Griesser M., Gustafsson L., Hansson B., Harris MP, Jiguet F., Kjellander P., Korpimäkki E., Krebs CH, Lens L, Linnell JCD, Low M, McAdam A, Margalida A., Merilä J., Moller AP, Nakagawa S., Nilsson JA, Nisbet ICT, van Noordwijk, Oro D, Pärt T, Pelletier F, Potti J, Pujol B, Réale D, Rockwell RF, Ropert-Coudert Y, Roulin A, Thébaud C, Sedinger JS, Swenson JE, Visser ME, Wanless S, Westneat DF, Wilson AJ, Zedrosser A. 2016. Solutions for archiving data in long-term studies: a reply to Whitlock et al. 2016. TREE 31: 85-87.
  • Reino, L.M., Borralho, R. & Arroyo, B. 2016. Influence of game crops on the distribution and productivity of red-legged partridges Alectoris rufa in Mediterranean woodlands. European Journal of Wildlife Research, 62: 609-617
  • Santangeli, A., Arroyo, B., Dicks L.V., Herzon, I., Kukkala, AS., Sutherland, W.J. & Moilanen A. 2016. Voluntary non-monetary approaches to nature conservation. Biological Conservation. 197: 209–214
  • Torres-Orozco, D., Arroyo, B., Pomarol, M. & Santangeli, A. (2016) From conservation traps to conservation solutions: lessons from intensively managed Montagu’s harriers in Catalonia. Animal Conservation 19: 436-443

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  • Cardador L., Brotons L., Mougeot F., Giralt D., Bota G., Pomarol M., Arroyo B. (2015) Conservation Traps and Long-Term Species Persistence in Human-Dominated Systems Conservation Letters 8: 456 – 462.
  • Jareño, D., Viñuela, J., Luque-Larena, J.J., Arroyo, L., Arroyo, B. & Mougeot, F. (2015) Factors associated with the colonization of agricultural areas by common voles Microtus arvalis in NW Spain. Biol. Invasions. 17: 2315.
  • Cardador L., Cáceres M.D., Giralt D., Bota G., Aquilué N., Arroyo B., Mougeot F., Cantero-Martínez C., Viladomiu L., Rosell J., Casas F., Estrada A., Álvaro-Fuentes J., Brotons L. (2015) Tools for exploring habitat suitability for steppe birds under land use change scenarios. Agriculture, Ecosystems and Environment 200: 119 – 125
  • Delibes-Mateos M., Viñuela J., Arroyo B. (2015) Game managers’ views on the release of farm-reared red-legged partridges in hunting estates within central Spain. Journal for Nature Conservation 26: 1 – 8.
  • Estrada A., Delibes-Mateos M., Caro J., Viñuela J., Díaz-Fernández S., Casas F., Arroyo B. (2015) Does small-game management benefit steppe birds of conservation concern? A field study in central Spain. Animal Conservation 18: 567-575
  • Tarjuelo R., Barja I., Morales M.B., Traba J. Bénitez-López A., Casas F., Arroyo B., Delgado M.P., Mougeot, F. (2015) Effects of human activity on physiological and behavioral responses of an endangered steppe bird. Behavioural Ecology, 26: 828-838.
  • Caro J., Delibes-Mateos M., Viñuela J., López-Lucero J.F., Arroyo B. (2015) Improving decision-making for sustainable hunting: regulatory mechanisms of hunting pressure in red-legged partridge. Sustainability Science 10: 479 – 489
  • Santangeli, A., Arroyo, B., Millon, A. & Bretagnolle, V. 2015. Identifying effective actions to guide volunteer-based and nation-wide conservation efforts for a ground-nesting farmland bird. J. Appl. Ecol. 52: 1082-1091.
  • Mills J.A., Teplitsky C., Arroyo B. Charmantier A., Becker P.H., Birkhead T.R., Bize P., Blumstein DT, Bonenfant C., Boutin S, Bushuev A., Cam E., Cockburn A., Côté SD, Coulson JC, Daunt F, Dingemanse NJ, Doligez B., Drummond H., Espie RHM, Festa-Bianchet M., Frentiu F., Fitzpatrick JW, Furness RW, Garant D., Gauthier G., Grant PR, Griesser M., Gustafsson L., Hansson B., Harris MP, Jiguet F., Kjellander P., Korpimäkki E., Krebs CH, Lens L, Linnell JCD, Low M, McAdam A, Margalida A., Merilä J., Moller AP, Nakagawa S., Nilsson JA, Nisbet ICT, van Noordwijk, Oro D, Pärt T, Pelletier F, Potti J, Pujol B, Réale D, Rockwell RF, Ropert-Coudert Y, Roulin A, Thébaud C, Sedinger JS, Swenson JE, Visser ME, Wanless S, Westneat DF, Wilson AJ, Zedrosser A. Archiving primary data: solutions for long-term studies. TREE 30: 581-589.
  • Delibes-Mateos, M., Mougeot, F., Arroyo, B. & Lambin, X. 2015. Hunted predator: charisma confounds. Science 349: 1294
  • Tarjuelo, R., Barja, I., Morales, M.B., Traba, J., Benitez, A., Casas, F., Arroyo, B. & Mougeot, F. 2015. Effects of weekend human activity on physiological stress and behavioral responses of an endangered steppe-bird. Behav. Ecol. 26: 828-838
  • Caro, J., Delibes-Mateos, M., Estrada, A. Borralho, R., Gordinho, L., Reino, L., Beja, P., & Arroyo, B. 2015. Effect of management for small-game on Mediterranean farmland birds. Bird Conserv. Intern. 25:166-181
  • Guzmán, J.L. & Arroyo, B. 2015. Predicting winter abundance of woodcock Scolopax rusticola using weather data: implications for hunting management. European J. Wild. Res. 61: 467-474
  • Limiñana, R., Arroyo, B., Terraube, J., McGrady, M. & Mougeot. F. 2015. Using satellite telemetry and environmental niche modelling to inform conservation targets for a long-distance migratory raptor in its wintering grounds. Oryx. 49: 329-337.
  • Madden, C.F., Arroyo, B. & Amar, A. 2015. A review of the impacts of crows, ravens and Eurasian Magpies on bird productivity and abundance. Ibis 157:1-16
  • Cardador, L., Cáceres, M., Giralt, D., Bota, G., Aquilué, N., Arroyo, B., Mougeot, F., Cantero-Martinez, C., Viladomiu, L., Rossell, J., Casas, F., Estrada, A., Alvaro-Fuentes, J. & Brotons, Ll. 2015. Tools for informing land use planning in agricultural systems: exploring habitat suitability for steppe birds under land-use change scenarios. Agric. Ecosyst. Env. 200: 119-125.

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  • Caro, J., Delibes-Mateos, M., Vicente, J. & Arroyo, B. 2014. A quantitative assessment of the release of farm-reared red-legged partridges (Alectoris rufa) for shooting in central Spain. E. J.Wildl. Res. 60: 919-926
  • Terraube, J. Guixé D. & Arroyo, B. 2014. Diet composition and foraging success in generalist predators: are specialist individuals better foragers? Basic and Applied Ecol. 15: 616-624
  • Santangeli, A., Di Minin, E. & Arroyo, B. 2014. Bridging the research implementation gap – Identifying cost-effective protection measures for Montagu’s harrier nests in Spanish farmlands. Biol. Cons. 177: 126-133
  • Delibes-Mateos, M., Giergiczny, M., Caro, J., Viñuela, J, Riera, P. & Arroyo, B. 2014. Does hunters´willingness-to-pay match the best hunting options for biodiversity conservation? A choice experiment application for partridge hunting in Spain. Biol. Cons. 177:36-42
  • Cardador, L., de Caceres, M., Bota, G., Giralt, D., Casas, F., Arroyo, B., Mougeot, F., Cantero, C., Moncunill, J., Butler, S. & Brotons, L. 2014. A resource-based modelling framework to assess the degree of habitat suitability for steppe birds in semiarid Mediterranean agricultural systems. PLOS One 9(3): e92790 (doi:10.1371/journal.pone.0092790)
  • Arroyo B., Leckie F., Amar, A., McCluskie, A. & Redpath, S. 2014. Ranging behaviour of hen harriers breeding in Special Protection Areas in Scotland. Bird Study 61: 48-55.
  • Jareño, D., Viñuela, J. Luque-Larena, J.J., Arroyo, L. Arroyo, B. & Mougeot, F. 2014. A comparison of methods for estimating common vole (Microtus arvalis) abundance in agricultural habitats. Ecological indicators 36: 111-119


  • Hobson, K. A., van Wilgenburg S.L., Guzmán, J.L. & Arroyo, B. 2013. Origins of juvenile Woodcock (Scolopax rusticola) harvested in Spain inferred from stable hydrogen isotope (δ2H) analyses of feathers. J. Ornithol. 154: 1087-1094
  • Talabante, C., Gómez, J., Aparicio, A. & Arroyo, B. 2013. Mixed clutches in Montagu´s harrier Circus pygargus nests: a maladaptive brood parasitism by galliform birds. Bird Study 60: 414-416.
  • Diaz-Fernandez, S., Arroyo, B. Viñuela, J., Patiño, I. & Riera, P. 2013. Market value of restocking and landscape management in red-legged partridge hunting. Wildlife Research 40: 336-343
  • Delibes-Mateos, M., Díaz-Fernández, S., Ferreras, P., Viñuela, J & Arroyo, B. 2013. The role of economic and social factors driving predator control in small game estates in central Spain. Ecology and Society. 18: 28.
  • Diaz-Fernandez, S., Arroyo, B., Casas, F., Haro, M., & Viñuela, J. 2013. Effect of management techniques on red-legged partridge abundance. PLOS One.
  • Luque-Larena, J.J., Mougeot, F., Viñuela, J., Jareño, D., Arroyo, L., Lambin, X., Arroyo, B.E. 2013. Recent large-scale range expansion and eruption of common vole (Microtus arvalis) outbreaks in agricultural plains of NW Spain. Basic and Applied Ecology 14: 432-441.
  • Arroyo, B. Mougeot, F. & Bretagnolle, V. 2013. Characteristics and functions of aerial displays of a semi-colonial raptor, the Montagu´s harrier. Journal Raptor. Res. 47: 185-196
  • Martinez-Padilla, J., Mougeot, F., García, J.T., Arroyo, B. & Bortolotti, G.R. 2013. Feather corticosterone levels and carotenoid-based coloration in common buzzard (Buteo buteo) nestlings. Journal Raptor Res. 47: 161-173
  • Mougeot, F., Gerrard, J., Dzus, E., Arroyo, B., Gerrard, P.N., Dzus, C. & Bortolotti, G. 2013. Population trends and productivity of Bald Eagles at Besnard Lake, Saskatchewan 1968-2010: density-dependent and climate effects. Journal Raptor Res. 47:96-107

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    • Diaz-Fernandez, S., Viñuela, J. & Arroyo, B. 2012. Harvest of Red-legged partridge in central Spain. J. Wild. Manag. 76:1354–1363
    • Estrada, A., & Arroyo, B. 2012. Occurrence vs abundance models: differences between species with varying aggregation patterns. Biol. Cons. 152: 37-45.
    • Arroyo, B., Delibes-Mateos, M., Diaz-Fernandez, S., & Viñuela, J. 2012. Hunting management in relation to profitability aims: red-legged partridge hunting in central Spain. E J Wildl Res. 58:847-855
    • Zuberogoitia, I., Arroyo, B., O’Donoghue, B.G., Zabala, J., Martínez, J.A., Martínez, J.E. & Murphy, S.G. 2012. Standing out from the crowd: are patagial wing tags a potential predator attraction for harriers (Circus spp.)? J. Orn. 153:985–989
    • Terraube, J., Mougeot, F, Bragin, A., Bragin, E. & Arroyo, B. 2012. Ecological factors influencing the breeding distribution and success of a nomadic specialist predator. Biodiv. Cons. 21: 1835-1852
    • Limiñana, R, García J.T., Guerrero A., González J.M., Lavedán J., Moreno J.D, Muñoz A-R, Palomares, L.E., Pinilla A., Ros G., Serrano C, Surroca M, Tena J. & Arroyo B. 2012.
    • Phylopatry and natal dispersal of Montagu’s harriers Circus pygargus breeding in Spain: a review of existing data. E J Wildl Res. 58: 549-555
    • Blanco-Aguiar, JA., Delibes-Mateos, M., Arroyo, B., Ferreras, P., Casas, F., Real, R., Vargas, JM., Villafuerte, R. & Viñuela, J. 2012. Is the interaction between rabbit hemorrhagic disease and hyperpredation by raptors a major cause of the red-legged partridge decline in Spain?. E J Wildl Res. 58: 433-439
    • Terraube, J., Mougeot, F., Cornulier, T., Verma, A., Gavrilov, A. & Arroyo, B. 2012. Broad wintering range and intercontinental migratory divide within a core population of the near-threatened pallid harrier. Divers. Dist. 18 : 401-409.
    • Limiñana, R., Soutullo, A., Arroyo, B. & Urios, V. 2012. Protected areas do not fulfil the wintering habitat needs of the tras-Saharan migratory Montagu’s harrier. Biol. Cons. 145:62-69
    • Garcia JT, Alda F, Terraube J, Mougeot F, Sternalski, A., Bretagnolle V & Arroyo B. 2011. The genetic effects of Holocene climatic change on steppe-associated fauna: demographic history, genetic structure and gene flow of in a raptor species. BMC Evol Biol. 11: 333 (doi:10.1186/1471-2148-11-333).
    • Terraube, J. & Arroyo, B.E. 2011. Factors influencing diet variation in a generalist predator across its range distribution. Biodiv. Cons. 20: 2111-2131
    • Guixé D., & Arroyo, B. 2011. Appropriateness of Special Protection areas for wide ranging species: the importance of scale and protecting foraging, not just nesting habitats. Animal Cons. 14: 391-399
    • Guzmán, J.L., Ferrand, Y. & Arroyo, B. 2011. Origin and migration of woodcock wintering in Spain. E. J. Wildl. Res. 57: 647-655
    • Limiñana, R., Arroyo, B., Surroca, M., Urios, V. & Reig-Ferrer, A. 2011. The influence of habitat on nest location and reproductive output of Montagu’s Harriers breeding in natural vegetation. J. Ornithol. 152: 557-565
    • Margalida, A., Moreno-Opo, R., Arroyo, B.E. & Arredondo, A. 2011. Reconciling the conservation of endangered species with economically important anthropogenic activities: interactions between cork exploitation and the cinereous vulture in Spain. Animal Cons. 14: 167-174.
    • Terraube, J., Arroyo, B.E, Madders, M. & Mougeot, F. 2011. Diet specialization and foraging efficiency under fluctuating food abundance in sympatric avian predators. Oikos 120: 234-244
    • Estrada, A., Arroyo, B. & Márquez, A.L. 2010. Possible changes in favourability areas for Montagu’s and hen harriers in Spain according to climate change. Ardeola 57(Especial): 123-128.
    • Milner-Gulland, E.J., Arroyo, B., Bellard, C., Blanchard, J., Bunnefeld, N., Delibes-Mateos, M., Edwards, C., Nuno, A., Palazy, L., Reljic, S., Riera, P., & Skrbinsek, T. 2010. New directions in management strategy evaluation through cross-fertilisation between fisheries science and terrestrial conservation. Biol. Lett. 6: 719-722.
    • Terraube, J., Arroyo, B.E., Mougeot, F., Katzner, T. & Bragin, E. 2010. Breeding biology of the Montagu’s harrier (Circus pygargus) in north-central Kazakhstan. J. Ornithol. 151:713-722.
    • Bertran, J., Margalida, A. & Arroyo, B. 2009. Agonistic behaviour and sexual conflict in atypical social reproductive groups: the case of bearded vulture polyandric trios. Ethology 115: 429-438.
    • Arroyo, B., Amar, A., Leckie, F., Buchannan, G., Wilson, J. & Redpath, S. 2009. Hunting habitat selection by hen harriers on moorland: implications for conservation. Biol. Cons. 142: 586-596
    • Anderson, B.J., Arroyo, B., Collingham, Y. C., Etheridge, B., Fernandez-de-Simon, J., Gillings, S., Gregory, R., Leckie, F., Thomas, C. D., Travis, J., Redpath, S.M. 2009. Using distribution models to test alternative hypotheses about a species’ environmental limits and recovery prospects. Biol. Cons. 142: 488-499
    • Terraube, J., Arroyo, B.E., Mougeot, F, Madders, M., Watson, J. & Bragin, EA. 2009. Breeding biology of the Pallid harrier Circus macrourus in North-Central Kazakhstan: implications for the conservation of a Near-Threatened species. Oryx 42: 1-9.
    • Amar, A., Amidon, A., Arroyo, B., Esselstyn, J.A. & Marshall, A.P. 2008. Population trends of the forest bird community on the pacific island of Rota, Mariana islands. Condor 110: 421-427.
    • Millon, A. Arroyo, B.E., & Bretagnolle, V. 2008. Variable but predictable prey availability affects predator breeding success: natural vs. experimental evidence. J. Zool. 275:349-358.
    • Leckie, F., Arroyo, B.E., Thirgood. S.J. & Redpath, S.M. 2008. Parental differences in brood provisioning by Hen Harriers Circus cyaneus. Bird Study 55: 209-215.
    • Hernández, M., Gonzalez, LM, Oria, J., Sánchez, R. & Arroyo B. 2008. Influence of contamination by organichlorine pesticides and polychlorinated biphenyls on the breeding of the Spanish Imperial Eagle (Aquila adalberti). Environ. Toxicol. Chem. 27: 433-441.
    • Amar, A., Arroyo, B., Meek E., Redpath, S. & Riley H. 2008. Influence of habitat on breeding performance of Hen Harriers in Orkney. Ibis 150: 400-404.
    • Arroyo, B.E., Bretagnolle, V. & Leroux, A.L.A. 2007. Interactive effects of food and age on breeding in the Montagu’s Harrier Circus pygargus. Ibis. 149: 806-813
    • Gonzalez, L.M., Arroyo, B.E., Margalida, A., Sánchez, R. & Oria, J. 2007. Can buffer zones have detrimental effects on the conservation of Spanish Imperial Eagles? Anim. Cons.10: 295-296.
    • Margalida, A., Arroyo, B.E., Bortolotti, G. & Bertran, J. 2006. Prolonged incubation in raptors: adaptive or nonadaptive behaviour? J. Raptor Res. 40 : 159-163
    • Bro, E., Arroyo, B. & Migot, P. 2006. Conflict between grey partridge hunting and hen harrier protection in France: a review. Wildlife Biology. 12: 279-293.
    • Redpath, SM, Leckie, F., Arroyo, B., Amar, A. & Thirgood, S. 2006. Compensating for the costs of polygyny in hen harriers Circus cyaneus. Behav. Ecol. Sociobiol. 60: 386-391. Mougeot, F. & Arroyo, B.E. 2006. Ultraviolet reflectance by the cere of raptors. Biol. Letters 2: 173-176.
    • Arroyo, B.E. & García, J.T. 2006. Diet composition influences annual breeding success of Montagu’s harriers Circus pygargus feeding on diverse prey. Bird Study 53: 73-78.
    • Gonzalez, L.M., Arroyo, B.E., Margalida, A., Sánchez, R. & Oria, J. 2006. Effect of human activities on the behaviour of breeding Spanish Imperial Eagles Aquila adalberti: management implications for the conservation of a threatened species. Anim. Cons. 9: 85-93.
    • Mougeot F., Arroyo B.E. & Bretagnolle, V. 2006. Paternity assurance responses to first-year and adult male territorial intrusions in a courtship feeding raptor. Anim. Behav. 71: 101-108.
    • Arroyo, B & Razin M. 2006. Effect of human activities on Bearded Vulture behaviour and breeding success in the French Pyrenees. Biol. Cons. 128: 276-284.
    • Morales, M.B., Bretagnolle, V. & Arroyo, B. 2005. Viability of the endangered Little Bustard Tetrax tetrax population of western France. Biodiv. Cons. 14: 3135-3150. Morales, M.B., García, J.T. & Arroyo, B. 2005. Can landscape composition changes predict spatial and annual variation of little bustard male abundance? Anim. Cons. 8: 167-174
    • Valkama, J., Korpimäki, E. Arroyo, B., Beja, P., Bretagnolle, V., Bro, E., Kenward, R., Mañosa, S., Redpath, S., Thirgood, S. & Viñuela, J. 2005. Birds of prey as limiting factors of gamebird populations in Europe: a review. Biol. Review 80: 171-203.
    • García, J.T. & Arroyo, B.E. 2005. Food-niche differentiation in sympatric Hen and Montagu’s harriers. Ibis 147: 144-154.
    • Arroyo, BE, Minguez, E, Palomares, L. & Pinilla, L. 2004. The timing and pattern of moult of flight feathers in the European Storm-petrel Hydrobates pelagicus in Atlantic and Mediterranean breeding areas. Ardeola 51: 365-373.
    • Amar, A., Arroyo, B., Redpath, S. & Thirgood S. 2004. Habitat variability predicts grouse predation rates by hen harriers. J. Appl. Ecol. 41: 305-314.
    • Redpath, S.M., Arroyo, B.E., Leckie, F., Bacon, P., Baifield, N., Gutierrez, R.J & Thirgood, S. 2004. Using decision modelling with stakeholders to reduce human-wildlife conflict: a raptor – grouse case study. Cons. Biol. 18: 350-359.
    • Arroyo, B.E. 2002. Sex biased nestling mortality in the Montagu’s harrier Circus pygargus. J. Avian Biol. 33: 455-460.
      García, J.T. & Arroyo, B.E. 2002. Intra and interspecific agonistic behaviour in sympatric harriers during the breeding season. Anim. Behav. 64: 77-84.
    • Redpath, S.M., Arroyo, B.E., Etheridge, B., Lekie, F.M., Bowman, K., Thirgood, S. 2002. Temperature and hen harrier productivity: from local mechanisms to geographical patterns. Ecography 25: 533-540.
    • Arroyo, B.E. 2002. Fledgling sex ratio variation and future reproduction probability in Montagu’s Harrier, Circus pygargus. Behav. Ecol. Sociobiol. 52: 109-116.
    • Arroyo, B.E., García, J.T. & Bretagnolle, V. 2002. Conservation of the Montagu’s Harrier (Circus pygargus) in agricultural areas. Anim. Conserv. 5: 283-290.
    • Arroyo B.E. & García, J.T. 2002. Alloparental care and kleptoparasitism in the semi-colonial Montagu’s Harrier Circus pygargus. Ibis 144: 676-679.
    • Jiguet, F., Jaulin, S. & Arroyo, B.E. 2002. Resource defence on exploded leks: do male little bustards, T. tetrax, control resources for females? Anim. Behav. 63: 899-905.
    • Morales, M.B., Jiguet, F. & Arroyo, B.E. 2001. Exploded leks: what bustards can teach us. Ardeola 48: 85-98.
      Arroyo, B.E., De Cornulier, Th. & Bretagnolle, V. 2002. Parental investment and parent-offspring conflicts during the post-fledging period in Montagu’s Harriers. Anim. Behav..63: 235-244.
    • García, J.T. & Arroyo, B.E. 2001. Effect of abiotic factors on reproduction in the centre and periphery of breeding ranges: a comparative analysis in sympatric harriers. Ecography 24: 393-402.
      Arroyo, BE, Mougeot, F & Bretagnolle, V. 2001. Colonial breeding and nest defence in Montagu’s harrier (Circus pygargus). Behav. Ecol. Sociobiol. 50: 109-115.
    • Mougeot, F., Arroyo, B.E. & Bretagnolle, V. 2001. Decoy presentations as a means to manipulate the risk of extrapair copulation: an experimental test of paternity assurance strategies in a semi-colonial raptor, the Montagu’s harrier Circus pygargus. Behav. Ecol. 12:1-7.
    • Jiguet, F., Arroyo, B. & Bretagnolle, V. 2000. Lek mating systems: a case study in the Little Bustard Tetrax tetrax. Behav. Process 51: 63-82.
    • Amar, A., Arroyo, B.E. & Bretagnolle, V. 2000. Post-fledging dependency and dispersal in hacked and wild Montagu’s Harriers Circus pygargus. Ibis 142: 21-28.
    • Arroyo, B.E., De Cornulier, Th. & Bretagnolle, V. 2000. Sex and age determination of short-eared owl nestlings. Condor 102:216-219.
    • Arroyo, B.E. & Bretagnolle, V. 1999. Breeding biology of the short-eared owl Asio flammeus in agricultural habitats in southwestern France. J. Raptor Res 33: 287-294.
    • Arroyo, B.E. & Bretagnolle, V. 1999. Field identification of Little Bustard males using plumage patterns. Ardeola 46: 53-60.
    • Wilson, J.D.; Morris, A.J.; Arroyo, B.E.; Clark, S.C.; Bradbury, R.B.1999. A review of the abundance and diversity of invertebrate and plant foods of granivorous birds in northern Europe in relation to agricultural change. Agric. Ecosyst. Env. 75: 13-30.
    • Arroyo, B.E. 1999. Copulatory behavior of semi-colonial Montagu’s Harrriers. Condor 101: 340-346.
    • Arroyo, B.E. 1998. Effect of diet on the reproductive success of Montagu’s Harrier Circus pygargus. Ibis 140: 690-693.
    • García, J.T. & Arroyo, B.E. 1998. Migratory movements of western European Montagu’s Harrier Circus pygargus: a review. Bird Study 45: 188-194.
    • Arroyo, B.E., Leroux, A.B.A. & Bretagnolle, V. 1998. Patterns of egg and clutch size variation in the Montagu’s Harrier. J. Raptor Res. 32: 136-142.
    • Arroyo, B.E. & García, J.T. 1997. Between-brood cannibalism in the Montagu’s Harrier. J Raptor Res 31: 286-287.
    • Palomares, L.E.; Arroyo, B.E.; Marchamalo, J.; Sainz, J.J. & Vesboren, B. 1997. Sex- and age-related biometric variation of Black-headed Gulls Larus ridibundus in Western European populations. Bird Study 44: 310-317.
    • Arroyo, B.E. 1997. Diet of Montagu’s Harrier Circus pygargus in central Spain: analysis of temporal and geographic variation. Ibis 139: 664-672.
    • Arroyo, B.E. & King, J.R. 1996. Age and sex differences in molt of the Montagu’s Harrier. J Raptor Res 30: 161-184.
    • Arroyo, B.E. 1996. Successful breeding by a first-year male Montagu’s Harrier. Bird Study 43: 383-384.
    • Arroyo, B.E. 1996. A possible case of polyandry in Montagu’s Harrier. J Raptor Res 30: 100-102.
    • Arroyo, B.E.; King, J.R. & Palomares, L.E. 1995. Observations on the ecology of Montagu’s and Marsh Harriers wintering in north-west Sénégal. Ostrich 66: 37-40.
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      • Brotons L., Bota G., Giralt D., Arroyo B., Mougeot F., Cantero C., Viladomiu L., 2010. “A step forward”. Economically viable farming and conservation of threatened bird species. Lynchos 3: 29-33
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      • Arroyo B., Pinilla A., Mougeot F., Cristal F., Guerrero A. 2009. Estudio de la ecología poblacional del aguilucho cenizo Circus pygargus en Extremadura. Servicio de Conservación de la Naturaleza y Espacios Naturales Protegidos, Junta de Extremadura pp 1-52.
      • Arroyo B., Pinilla A., Mougeot F., Cristal F., Guerrero A. 2009.Estu dio de la ecología poblacional del aguilucho cenizo Circus pygargus en Extremadura. Servicio de Conservación de la Naturaleza y Espacios Naturales Protegidos, Junta de Extremadura pp 1-52.

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